I looked at the output of listkeys like you said. It looks like you are right, it is just on the server. I checked the different characters and they were all the same.

If I type listkeys there are 5

KilledTroll
killed_surtling
defeated_queen
defeated_eikthyr
defeated_hive


Which is, honestly all sorts of messed up!

Every boss was spawned and killed except the queen and hive; so it looks like at some point a player killed those bosses and I didn't notice and I saved it to the server

But The Elder Key is missing for some reason, and also Moder

I think the hive boss might be messing with the logic...

All sorts of weird...